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Sequence-directed RNA remodeling within a topologically complex RNP substrate

bioRxiv, ISSN: 2692-8205
2022
  • 4
    Citations
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Metrics Details

  • Citations
    4
    • Citation Indexes
      4
      • CrossRef
        4

Article Description

DEAD-box ATPases are ubiquitous enzymes essential in all aspects of RNA biology. However, the limited in vitro catalytic activities described for these enzymes is at odds with their complex cellular roles, most notably in driving large-scale RNA remodeling steps during the assembly of ribonucleoproteins (RNPs). We describe cryo-EM structures of 60S ribosomal biogenesis intermediates that reveal how context-specific RNA unwinding by the DEAD-box ATPase Spb4 results in extensive, sequence-directed remodeling of rRNA secondary structure. Multiple cis and trans interactions stabilize a post-catalytic, high-energy intermediate that drives the organization of the root helix structure within rRNA domain IV. This mechanism explains how limited strand separation by DEAD-box ATPases is leveraged to provide non-equilibrium directionality and ensure efficient and accurate RNP assembly.

Bibliographic Details

Victor Emmanuel Cruz; Kamil Sekulski; Nagesh Peddada; Sahana Balasubramanian; Christine S. Weirich; Jan P. Erzberger; Carolin Sailer; Florian Stengel

Cold Spring Harbor Laboratory

Biochemistry, Genetics and Molecular Biology; Agricultural and Biological Sciences; Immunology and Microbiology; Neuroscience; Pharmacology, Toxicology and Pharmaceutics

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